2011 Annual Checklist DVD front cover
This release of the Catalogue of Life contains contributions from 143 databases with information on 1,578,063 species, 146,277 infraspecific taxa and also includes 1,277,567 synonyms and 390,260 common names covering the following groups:

Viruses • Viruses and Subviral agents from ICTV_MSL

Bacteria and Archaea from BIOS

Chromista • Chromistan fungi from Species Fungorum UPDATED!

Protozoa • Major groups from ITIS Regional, • Ciliates from CilCat, • Polycystines from WoRMS Polycystina UPDATED!, • Protozoan fungi from Species Fungorum UPDATED! and Trichomycetes UPDATED! database • Microsporidians from Microsporidia NEW! • Slime moulds from Nomen.eumycetozoa.com UPDATED!

Fungi • Various taxa in whole or in part from CABI Bioservices databases (Species Fungorum UPDATED!, Phyllachorales UPDATED!, Rhytismatales UPDATED!, Saccharomycetes UPDATED! and Zygomycetes UPDATED! databases) and from three other databases covering Xylariaceae UPDATED!, Glomeromycota UPDATED!, Trichomycetes UPDATED!, Dothideomycetes UPDATED! • Lichens from LIAS

Plantae (Plants) • Mosses from MOST • Liverworts and hornworts from ELPT • Ferns and fern-allies from World Ferns NEW! • Conifers from Conifer Database UPDATED! • Cycads and 5 flowering plant families from IOPI-GPC, 99 families from WCSP, 280 families from World Plants NEW! and 14 families from SCaDs NEW! • Plus individual flowering plants families from AnnonBase UPDATED!, Brassicaceae, ChenoBase UPDATED!, Droseraceae Database, EbenaBase UPDATED!, GCC UPDATED!, ILDIS, ILDIS for i4Life NEW!, LecyPages, LHD, MELnet, RJB Geranium, Solanaceae Source, Umbellifers.

Animalia (Animals) • Marine groups from URMO, ITIS Global UPDATED!, Hexacorals, ETI WBD (Euphausiacea), WoRMS: WoRMS Asteroidea UPDATED!, WoRMS Bochusacea UPDATED!, WoRMS Brachiopoda UPDATED!, WoRMS Brachypoda UPDATED!, WoRMS Brachyura UPDATED!, WoRMS Bryozoa UPDATED!, WoRMS Cestoda UPDATED!, WoRMS Chaetognatha UPDATED!, WoRMS Cumacea UPDATED!, WoRMS Echinoidea UPDATED!, WoRMS Gastrotricha UPDATED!, WoRMS Gnathostomulida UPDATED!, WoRMS Holothuroidea UPDATED!, WoRMS Hydrozoa UPDATED!, WoRMS Isopoda UPDATED!, WoRMS Leptostraca UPDATED!, WoRMS Monogenea UPDATED!, WoRMS Mystacocarida UPDATED!, WoRMS Myxozoa UPDATED!, WoRMS Nemertea UPDATED!, WoRMS Oligochaeta UPDATED!, WoRMS Ophiuroidea UPDATED!, WoRMS Phoronida UPDATED!, WoRMS Placozoa UPDATED!, WoRMS Polychaeta UPDATED!, WoRMS Polycystina UPDATED!, WoRMS Porifera UPDATED!, WoRMS Priapulida UPDATED!, WoRMS Proseriata-Kalyptorhynchia UPDATED!, WoRMS Remipedia UPDATED!, WoRMS Scaphopoda UPDATED!, WoRMS Tanaidacea UPDATED!, WoRMS Tantulocarida UPDATED!, WoRMS Thermosbaenacea UPDATED!, WoRMS Trematoda UPDATED!, WoRMS Xenoturbellida UPDATED! • Rotifers, mayflies, freshwater hairworms, planarians, water fleas and aquatic mites from FADA databases: FADA Rotifera, FADA Ephemeroptera, FADA Nematomorpha, FADA Turbellaria, FADA Cladocera NEW! & FADA Halacaridae NEW! • Entoprocts, water bears from ITIS Global UPDATED! • Spiders, scorpions, ticks & mites from SpidCat via ITIS, SalticidDB , ITIS Global UPDATED!, TicksBase, SpmWeb BdelloideaBase UPDATED! & Mites GSDs: OlogamasidBase, PhytoseiidBase, RhodacaridBase & TenuipalpidBase • Diplopods, centipedes, pauropods and symphylans from SysMyr UPDATED! & ChiloBase • Dragonflies and damselflies from Odonata database • Stoneflies from PlecopteraSF • Cockroaches from BlattodeaSF • Praying mantids from MantodeaSF • Stick and leaf insects from PhasmidaSF • Grasshoppers, locusts, katydids and crickets from OrthopteraSF • Webspinners from EmbiopteraSF • Bark & parasitic lices from PsocodeaSF • Some groups of true bugs from ScaleNet, FLOW, COOL, Psyllist, AphidSF , MBB, 3i Cicadellinae, 3i Typhlocybinae, MOWD, CoreoideaSF, The White-Files NEW!, Tessaratomidae Database NEW! & Lace Bugs Database NEW!• Twisted-wing parasites from Strepsiptera Database • Lacewings, antlions, owlflies, fishflies, dobsonflies & snakeflies from LDL Neuropterida UPDATED! • Some beetle groups from the Scarabs UPDATED!, TITAN, WTaxa, CarabCat NEW! & ITIS Global UPDATED! • Fleas from Parhost • Flies, mosquitoes, bots, midges and gnats from Systema Dipterorum, CCW UPDATED! & CIPA • Butterflies and moths from LepIndex, GloBIS (GART) UPDATED!, Tineidae NHM, Global Gracillariidae • Bees & wasps from ITIS Bees, Taxapad Ichneumonoidea, UCD, ZOBODAT Vespoidea & HymIS UPDATED!• Molluscs from WoRMS Mollusca UPDATED!, FADA Bivalvia, MolluscaFW & AFD (Pulmonata) • Fishes from FishBase UPDATED! • Reptiles from ReptileDB UPDATED! • Amphibians, birds and mammals from ITIS Global UPDATED!

PLUS additional species of many groups from ITIS Regional, NZIB and CoL China


The Catalogue of Life Editors

The Catalogue of Life combines the outputs of the Species 2000 and the ITIS programmes. Assembly and publication of the Catalogue of Life is managed by Yuri Roskov in Reading, working with colleagues around the world.

Yuri Roskov, Catalogue of Life Executive Editor at the University of Reading, UK.

Thomas Kunze, Catalogue of Life Editorial Assistant at the CoL Secretariat.

Luisa Abucay and Luvie Paglinawan (until July 2013), Catalogue of Life Database Managers at the CoL Philippines Office, FIN, Los Baños, Philippines.

Thomas Orrell, ITIS Deputy Director, ITIS Secretariat, Smithsonian Institution, Washington DC, USA.

Alastair Culham, PI for i4Life EC project at the University of Reading, UK.

Nicolas Bailly, Taxonomic Editorial Advisor for Marine sectors at the CoL Philippines Office, FIN, Los Baños, Philippines.

Paul Kirk, Taxonomic Editorial Advisor for Fungi and Chromista sectors at the Royal Botanic Gardens, Kew, Richmond, UK.

Thierry Bourgoin, Taxonomic Editorial Advisor for Insects at the Museum National d'Histoire Naturelle, Paris, France.

Guy Baillargeon, Taxonomic Editorial Advisor for Plants, Canada.

Wim Decock, Taxonomic Editorial Advisor for marine taxa (WoRMS) at the Flanders Marine Data Centre, VLIZ, Oostende, Belgium

Aaike De Wever, Taxonomic Editorial Advisor for freshwater taxa (FADA) at the Royal Belgian Institute of Natural Sciences, Brussels, Belgium.

Viktoras Didžiulis (until November 2013), editorial transformations of World Ferns, World Plants and WoRMS data at the University of Reading, UK.


The Catalogue of Life Assembly Team

Luisa Abucay, Catalogue of Life Database Manager at the CoL Philippines Office, FIN, Los Baños, Philippines.

Viktoras Didžiulis, Production Systems Manager & Developer at the CoL Secretariat (until November 2013), University of Reading, UK.


Annual Checklist Software Development

Wouter Addink, Leader of the Annual Checklist Software Development and Software Production at Naturalis-ETI, Leiden, The Netherlands.

Ruud Altenburg, author of Base Schema Conversion Software at Naturalis-ETI, Leiden, The Netherlands.

Ayco Holleman, Annual Checklist Assembly ICT Infrastructure Development at Naturalis-ETI, Leiden, The Netherlands.

Gideon Gijswijt, Annual Checklist DVD Master and Installation Development at Naturalis-ETI, Leiden, The Netherlands.

Viktoras Didžiulis, author of the CoL Workbench and data assembly workflow design as former Systems Manager at the University of Reading, UK.


Dennis Seijts, former Annual Checklist Interface Development at ETI Bioinformatics, Amsterdam, The Netherlands.

Núria Torrescasana Aloy, former author of the Annual Checklist Software at ETI BioInformatics, Amsterdam, the Netherlands.

Jorrit van Hertum, former author of the Annual Checklist Software at ETI BioInformatics, Amsterdam, the Netherlands.

Richard White, former author of the Taxon Matcher Software at Cardiff University, UK.

Paul Klinkenberg, Annual Checklist booklet design and DVD production for Naturalis-ETI, Leiden, The Netherlands.


The cover picture of Cystopteris fragilis is kindly provided by Michael Hassler.